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Stork migration

Paqe and Ana, two stork sisters from Kosovo: they left by different roads and were 27 metres apart again in Sudan

In July 2017, in a nest in the village of Prelluzhë in Kosovo, two stork sisters, Paqe (Albanian for “peace”) and Ana, were fitted with 50-gram loggers that recorded their position every five minutes. They left for Africa four days apart, by different roads. Six weeks later, 3,284 km from home in eastern Sudan, they were 27 metres apart. The next summer, Paqe spent a month in Romania.

How far apart they were, every day

The gap between the threads is the closest the storks came to each other each day, on a scale from a few metres to a thousand kilometres. The bright ticks are the days they were less than a kilometre apart. Touch the threads to see the day.

The distance between Paqe and Ana, day by day, July 2017 to January 2019AugSepOctNovDecJan 2018FebMarAprMayJunJulAugSepOctNovDecJan 2019PaqeAnaPrelluzhë, Kosovo27 m · eastern Sudan390 m · northern KenyaAna dies in SyriaPaqe · summer in Romanialast fix · TanzaniaThe distance between Paqe and Ana, day by day, July 2017 to January 2019AugSepOctNovDecJan 2018FebMarAprMayJunJulAugSepOctNovDecJan 2019PaqeAnaPrelluzhë, Kosovo27 m · eastern Sudan390 m · northern KenyaAna dies in SyriaPaqe · summer in Romanialast fix · Tanzania

Data: the LifeTrack White Stork Kosova project, published on Movebank: 122,097 valid positions for Paqe, 74,076 for Ana.

A nest in Prelluzhë

Prelluzhë is a village in Vushtrri municipality, north of Pristina. Paqe and Ana hatched no later than mid-May 2017, in a nest with three chicks; Paqe was the elder of the two that were tagged. On 5 July the German ornithologist Wolfgang Fiedler and the students on a course fitted each with a 50-gram solar-powered logger that sends its positions over the mobile phone network.

“Paqe” is Albanian for peace, from the Latin pax. On 20 July, on the same morning and an hour and 40 minutes apart, the sisters went more than a kilometre from the nest for the first time.

They left four days apart

Ana left first: by 11:15 on 15 August 2017 she was already more than 50 kilometres from the nest, and that night she slept in Serbia. Paqe was that far from the nest by 12:35 on 19 August 2017, four days later. Ana went through Serbia and Bulgaria, Paqe through Serbia, North Macedonia and Bulgaria; both then crossed a corner of Greece into Turkey.

Storks travel by soaring: they climb in columns of warm air rising from hot ground, then glide to the next one. No such columns form over the sea, so they avoid water. Ana crossed the Sea of Marmara all the same, on 20 August 2017: between 09:30 and 11:55, most of her positions are over the water.

Through Turkey, Syria, Lebanon or Jordan, Israel and Sinai to Sudan

From Turkey both went south through the Levant, away from the coast: Ana through Syria and Lebanon, Paqe through Syria, past Damascus, and Jordan. They crossed southern Israel into Sinai, flew over the Gulf of Suez and followed the Nile south. Ana reached Sudan on 29 August, Paqe on 31 August, on a day she flew 463 km.

Six weeks later, in the same region

In September both stopped in eastern Sudan, in Kassala state, each arriving by her own road. On 22 September 2017, at about half past two in the afternoon, Paqe landed beside Ana; at 14:35 they were 45 metres apart. On 27 September 2017, at 11:20, their loggers were 27 metres apart, both on the ground.

In September and October they were less than a kilometre apart on 10 days, but on most days they were tens of kilometres apart: in the same region, not together. To see whether such closeness also turns up by chance, Ana's track was shifted by one to fifteen days, forwards and back. In the shifted versions the two come within a kilometre on at most 5 days, not 10, and never within 50 metres.

Whether they recognised each other cannot be known. The loggers say only where each one was.

Twenty-seven metres

This is the place, in a 2016 satellite image: a plain in Kassala state, some 70 km north-north-west of the town of New Halfa. The orange dots are Paqe's positions on 27 September 2017, the blue ones Ana's.

Winter, over the Ethiopian mountains and on to Tanzania

In November Paqe crossed the Ethiopian highlands. On 20 November 2017 she climbed to 4,147 metres above sea level, the highest reliable height her logger recorded, 2,384 metres above the mountains. Ana went through Eritrea and Ethiopia; in December she too climbed above 4,500 metres.

On 21 December 2017, in northern Kenya, the sisters were close again: 390 metres. After New Year both were in Tanzania, but in different places; they stayed there until the third week of March.

North, in spring

Around 22 March 2018 both set off north. Paqe flew almost without a break as far as Sudan and stayed there until mid-April; Ana spent two weeks in southern Kenya and set off again on 9 April. Further north their roads ran nearly parallel. In Israel and on the Lebanese border, between 29 April and 13 May, the smallest distance between them each day was between 21 km and 72 km. Then Paqe flew on and by 14 May was in Turkey, while Ana stayed a few days in Lebanon and went on into Syria.

Ana, in northern Syria

On 26 May 2018 Ana stopped near Afrin, in north-western Syria. Her logger did not move again. The project's record says she died on 27 May and that the logger's motion sensor “shows hunting or predation event”. The logger went on sending positions from the same spot; the published data end on 31 July.

Storks are shot along this route. In Lebanon, of 77 rings from white storks found and reported, 75.3% came from birds confirmed as shot, according to a 2021 study.

Paqe's summer, in Romania

Paqe was just over a year old. She spent May and June in Turkey, then flew up through Bulgaria and on 11 July 2018 crossed the Danube into Romania's Călărași county near Ciocănești; she spent her first night near Nicolae Bălcescu. She stayed 33 days.

In the first days she flew north, to around Ploiești, then west into Dâmbovița county.

Six days at the rubbish dump outside Bucharest

From 16 to 21 July Paqe stayed on the western edge of Bucharest, near the village of Rudeni. Between 7 a.m. and 7 p.m., 39% of her positions fall inside the outline OpenStreetMap draws for the IRIDEX landfill at Chiajna. At night she slept less than a kilometre away. Storks feed at rubbish dumps: in Flack and colleagues' 2016 study, four of the six storks from south-west Germany that lived at least five months spent the winter at dumps in Morocco.

Two and a half weeks near the Danube

On 22 July she headed south, and from 25 July to 11 August she stayed on the plain by the Danube, between Giurgiu and Teleorman counties, usually about 10 kilometres from the river: around the villages of Găujani, Pietrișu, Izvoru, Bujoru and Cervenia. On 12 August she crossed the Danube back into Bulgaria and headed south.

The second autumn

On 26 August 2018, over the desert of southern Egypt, she flew 515 km in a single day, the longest of her life. She reached Sudan on 27 August, went through South Sudan and Ethiopia, spent December in Kenya and on 11 January 2019 entered Tanzania.

The last day

On 22 January 2019, in central Tanzania, Paqe flew 64 km. At 15:00 she climbed to 2,257 metres. The last valid position is from 22:00, at the place where she had settled for the night. The project's record says only that she died that day, of unknown causes.

Her logger followed her for 567 days and recorded 122,097 valid positions and, added up, the distances between them come to more than 46,000 km, more than once round the Earth. She passed through 20 countries and territories.

The other six storks from Kosovo

From 2016 to 2018 the project fitted loggers to eight stork chicks from three nests in Kosovo. None was tracked to the age of two. Seven died, and one vanished:

  • Alban, Agron and Besi: three brothers from one nest near Lipjan, tagged in 2016. Alban died in western Turkey on 4 September 2016, and his logger was then carried by a person for a long time. Besi and Agron were “presumably hunted” in Sudan in November and December 2016; Agron's logger was taken into a house.
  • Lean, Lumbardha and Drin: three siblings from the village of Pobërgjë, tagged in 2018. Lumbardha died in Egypt around 2 September 2018, and her logger was found two years later. Drin died in January 2019 in Tanzania's Mbeya region. Lean vanished on 4 February 2019 near Dodoma, Tanzania; until then his few positions showed normal movement.

Eight birds are too few to say how long storks from Kosovo live. A larger study, of 70 stork chicks tagged in eight countries, found 21 still alive after a year, or 30%. Of the ten from Greece, the population nearest to Kosovo, two were alive.

A day in Paqe's life

Pick a day between 5 July 2017 and 22 January 2019: your birthday, an anniversary, a day you remember. See where Paqe was, how far she flew, and how she climbed and sank that day over the ground beneath her.

The eight storks from Kosovo

StorkNestTaggedDeath or loss (per the record)What the project's record says
PaqePrelluzhë5 July 201722 January 2019Died of unknown causes on 22 January 2019 in Tanzania.
AnaPrelluzhë5 July 201727 May 2018Died in Syria on 27 May 2018. “ACC shows hunting or predation event. logger continued sending until October, but no chance to retrieve the logger”
AlbanLipjan23 June 20164 September 2016“died in W Turkey 2016-09-04 afternoon in fields. Logger then was transported by man for a long time. No obvious reason for death.”
AgronLipjan23 June 20166 December 2016“presumably hunted 2016-12-06 in SW Sudan. Logger was brought to a house.”
BesiLipjan23 June 20167 November 2016“presumably hunted 2016-11-07 in S of Sudan. Logger close to settlement.”
LumbardhaPobërgjë20 June 20182 September 2018Dead of unknown causes since about 2 September 2018 in Egypt. “Logger found after 2 years!”
DrinPobërgjë20 June 20189 January 2019Died on 9 January 2019 in Mbeya, Tanzania. “Logger not moving for many months. Before, too few coordinates to decide about reason of death”
LeanPobërgjë20 June 20184 February 2019Vanished on 4 February 2019 near Dodoma, Tanzania. “GPS shows normal movement (although few coordinates) before signal was lost.”

How it is calculated

The positions come from the LifeTrack White Stork Kosova dataset, published in the Movebank Data Repository under CC0. The 50-gram e-obs loggers take a GPS fix every five minutes by day and less often at night. The page uses only positions marked valid and with a 3D fix.

The distance between the sisters is measured between their positions taken no more than 90 seconds apart, using only positions whose accuracy, as estimated by the logger, is 20 metres or better (half of all positions are accurate to 4.4 metres or better). The chance test shifts Ana's track by 1 to 15 days forwards and back and counts, in the same Sudan window (7 September to 14 October 2017), the days on which the two come within a kilometre and the positions within 50 metres.

Distance flown adds up the segments between successive positions that are longer than 200 metres or flown at more than 2 m/s, so it does not count the GPS jitter of a bird standing still. Straight lines between positions cut the real path slightly short, so the figure is a lower bound.

Height above sea level is the GPS height above the ellipsoid minus the EGM96 geoid. The ground beneath the bird comes from the Copernicus GLO-90 model (90 metres); at rest, Paqe's and Ana's positions sit on average 2.4 metres above it, which shows the two agree. Left out of the heights are isolated positions that jump more than 400 metres from both neighbours in the same direction, and those that would need a vertical speed above 8 m/s.

The country comes from Natural Earth borders; the place is the nearest one in GeoNames. Times are the local time of the country the bird was in.

The project's records are quoted from the published table (in English). What happened to Paqe on 22 January 2019 is not known.

Sources

  1. Maxhuni Q, Gashi A, Hoxha L, Wolf G, Fiedler W (2021). Data from: Study “LifeTrack White Stork Kosova”. Movebank Data Repository, CC0. doi.org
  2. Kays R et al. (2021). The Movebank system for studying global animal movement and demography. Methods in Ecology and Evolution. doi.org
  3. Flack A et al. (2016). Costs of migratory decisions: a comparison across eight white stork populations. Science Advances 2: e1500931 (table 1). doi.org
  4. Raine AF et al. (2021). The international dimension of illegal bird hunting in Lebanon. Sandgrouse 43. www.spnl.org
  5. NASA Earth Observatory. Blue Marble Next Generation, July 2004 (public domain). visibleearth.nasa.gov
  6. NOAA NCEI. ETOPO1 Global Relief Model, via ERDDAP. upwell.pfeg.noaa.gov
  7. Copernicus DEM GLO-90 (ESA), © DLR e.V. 2010–2014 and © Airbus Defence and Space GmbH 2014–2018, provided under the Copernicus programme. registry.opendata.aws
  8. NGA. EGM96 geoid model, 15′ grid (PROJ). cdn.proj.org
  9. EOX IT Services. Sentinel-2 cloudless 2016 (contains modified Copernicus Sentinel data 2016), CC BY 4.0. s2maps.eu
  10. OpenStreetMap, outline “IRIDEX - Chiajna” (way 334645076), ODbL. www.openstreetmap.org
  11. GeoNames, places with more than 500 inhabitants, CC BY 4.0. www.geonames.org
  12. Natural Earth: borders, rivers and lakes (public domain). www.naturalearthdata.com
  13. Wiktionary: Albanian “paqe”. en.wiktionary.org