Figures come from scientific papers, databases and reference guides; calculations made for this page are labelled separately. The list says what each source was used for. Where sources give different values, the range is printed: the 12,000–15,000 km circuit, the 1,500–2,000 km Sahara, the 50–60 mm wingspan.
No butterfly on this map carried a GPS tracker. The route is reconstructed from several kinds of evidence, with different uncertainties — the table below sets them side by side.
Two things stay hard to pin down: how long a wild adult lives, which is hard to measure directly, and the exact route of the butterflies reaching Romania. The page says so plainly next to each.
The page's own calculations — the Bucharest–Sahel distance, the June share, how many butterflies span the Sahara — are automatically verified at every release of the page.
How do you track a butterfly without GPS?
- Wing isotopes tell where it was born
- Pollen visited plants give geographical clues
- DNA shows the population and the origin
- Wind models reconstruct the feasible route
The claim, the evidence, its strength
| Claim | Evidence | Strength and limit |
|---|
| Europe–sub-Saharan Africa, autumn | isotopes and observations | direct support for some individuals |
|---|
| Sub-Saharan Africa–Mediterranean, spring | hydrogen isotopes | strong evidence for many sampled individuals |
|---|
| Monthly tropical breeding zones | fieldwork, monitoring and niche modelling | regional and monthly, not individual routes |
|---|
| Eastern Mediterranean–East Africa | dual hydrogen and strontium isotopes | first direct evidence, 2024 |
|---|
| The crossing to French Guiana | wind, genes, pollen, isotopes and modelling | solid reconstruction of a rare event |
|---|
| Romanian seasonality | GBIF records | shaped by observer effort |
|---|
The circuit and the flight
- primary study Stefanescu, C. et al. (2013). Multi-generational long-distance migration of insects: studying the painted lady butterfly in the Western Palaearctic. Ecography 36(4). doi:10.1111/j.1600-0587.2012.07738.x — the older six-generation reconstruction; the circuit of up to 15,000 km; the radar-measured speeds and heights (accepted manuscript read)
- primary study Stefanescu, Soto, Talavera, Vila, Hobson (2016). Long-distance autumn migration across the Sahara by painted lady butterflies. Biology Letters 12: 20160561. doi:10.1098/rsbl.2016.0561 — the flights of over 4,000 km from Europe to the savannah, proven with isotopes
- primary study Talavera, Bataille, Benyamini, Gascoigne-Pees, Vila (2018). Round-trip across the Sahara. Biology Letters 14: 20180274. doi:10.1098/rsbl.2018.0274 — the spring return, the circuit of about 12,000 km, winter in the Afrotropics
- primary study Talavera et al. (2023). The Afrotropical breeding grounds of the Palearctic-African migratory painted lady butterflies. PNAS 120(16). doi:10.1073/pnas.2218280120 — the tropical breeding grounds, up to ten generations a year, the monthly maps behind the zones drawn here
- primary study Reich et al. (2024). Trans-Saharan migratory patterns in Vanessa cardui and evidence for a southward leapfrog migration. iScience 27: 111342. doi:10.1016/j.isci.2024.111342 — broad-front migration, moderate connectivity, leapfrogging, the eastern-Mediterranean–East Africa link, the 8–10 overlapping-generations framing; 3–6-week lifespan, over 1,000 eggs
- primary study Reich et al. (2025). Isotope geolocation and population genomics in Vanessa cardui: short- and long-distance migrants are genetically undifferentiated. PNAS Nexus 4: pgae586. doi:10.1093/pnasnexus/pgae586 — short- and long-distance migrants, wintering on both sides of the Sahara, the lack of genetic differentiation
- primary study Suchan et al. (2024). A trans-oceanic flight of over 4,200 km by painted lady butterflies. Nature Communications 15: 5205. doi:10.1038/s41467-024-49079-2 — the Atlantic crossing, in 5–8 days, on wind and gliding; the energy models' limits
- primary study García-Berro et al. (2025). A north-south hemispheric migratory divide in the butterfly Vanessa cardui. Nature Communications 16: 11341. doi:10.1038/s41467-025-67185-7 — the hemispheric migratory divide, the ~9 Mb inversion, the equatorial contact zone (the 'further south' box)
The butterfly's life
- guide Myers et al. Vanessa cardui. Animal Diversity Web, University of Michigan. animaldiversity.org — development, host plants, the migration calendar, laboratory life
- guide Butterfly Conservation. Painted Lady. butterfly-conservation.org — the 50–56 mm wingspan, thistles as the preferred plant
- guide Beneš, Dvořák. Vanessa cardui, identification key card. lepidoptera.cz. lepidoptera.cz — the 55–60 mm wingspan
- database Catalogue of Life, via GBIF: the Romanian name Fluturele de scaieți. gbif.org — the species' Romanian name
Romania and the data
- database GBIF, Vanessa cardui records from Romania (exact query: taxon 4299368, country RO), accessed 23 September 2026. gbif.org/occurrence/search — 961 records: months, years, sources; 15 without a month