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130,000 years of giants and people
Mammoths, giant sloths, giant kangaroos: the animals that vanished after people arrived
Without the extinctions of the last 130,000 years, 329 species of land mammal heavier than 44 kg would be alive today. 175 are gone. Scroll, and watch where they disappear and when: on every continent but Africa, most of the losses come after the first modern humans.
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130,000 years ago
A planet of giants
The ochre shows how many kinds of large land mammal, over 44 kg, would live in each place today if none had died out since this moment. The brightest land holds around 45. Worldwide there are 329 species.
The firelight is people. Here it burns only in Africa, where our species began. The faint embers in Europe and Asia are Neanderthals and other human species.
Africa
Where people have always been
Africa lost 16 of its 82 giant species. Elephants, rhinos, hippos, giraffes and buffalo still live there, and nowhere else do so many giant species still live side by side.
Humans and their ancestors lived alongside these animals for over two million years. One explanation is that the animals had time to learn to fear them. Not everyone agrees: Tyler Faith (2014) finds Africa’s losses driven by grasslands changing, with “little reason” to blame people.
Australia and New Guinea · 48,000 years ago
The first new continent
People were in Australia by about 48,000 years ago, most researchers agree; tools at Madjedbebe have been dated to 65,000, a date others reject. A model of the fossil dates puts a peak of extinctions at 42,100 years ago.
Of the 26 giant mammals that lived here, 25 are gone: Diprotodon, a 2.7-tonne marsupial, kangaroos two metres tall, a marsupial lion. A flightless bird taller than a person, kin to ducks and geese, and a lizard several metres long went too; the ochre counts only mammals.
How soon is soon? In south-west Australia, dung fungus that thrives on big herbivores collapses between 45,000 and 43,100 years ago, “within 4,000 years” of people. In north-east Queensland some giants lasted until after 40,000 years ago, and some researchers think most species were gone before people came.
Eurasia · 45,000 years ago
Eurasia: a slow fade
Our species was in Bulgaria before 45,000 years ago. In Romania, the jaw from Peștera cu Oase is 37,000 to 42,000 years old. By about 40,000 years ago the Neanderthals were gone: they were large mammals too, though they are not counted here.
Then the giants went one by one, over tens of thousands of years: the cave bear around 30,000 years ago, the cave lion and the woolly rhinoceros around 14,000, the woolly mammoth on the mainland around 10,700, the giant deer after 7,700. Eurasia lost 66 of 147 species (45%).
Here the warming at the end of the Ice Age clearly mattered. Forest replaced the mammoth steppe, and each species’ range shrank on its own schedule. People were one more pressure; how much is still argued.
The Americas · 16,000 years ago
The Americas: a few thousand years
People were at Cooper’s Ferry in Idaho by about 16,000 years ago and at Monte Verde in Chile by 14,200. Footprints at White Sands, New Mexico, are 21,000 to 23,000 years old.
The last dated fossils of 16 large North American genera fall between 13,800 and 11,400 years ago; in South America the record collapses between 12,900 and 11,600. North America lost 50 of 70 giant species, South America 50 of 65: mammoths, mastodons, ground sloths as heavy as elephants, armoured glyptodonts, horses, camels, sabre-toothed cats.
If the White Sands footprints are right, people and these animals shared the continent for several thousand years before the collapse. That fits “after people arrived”, and argues against a sudden slaughter.
The Caribbean
The Caribbean: a test that wobbles
Ground sloths outlived their mainland cousins on Cuba and Hispaniola by thousands of years, which is what you would expect if people were the threat: no people, no extinction.
The newest dating of the first settlements, though, puts people on Cuba 5,400 to 4,700 years ago and on Hispaniola later still. On that timeline most of the last dated sloth bones are older than the first people. With so few dated bones this cannot settle it, and the page does not count it as a clean case.
Two exceptions
Where mammoths died without people
On St Paul Island in the Bering Sea, mammoths lived until 5,600 years ago and died as the island shrank and its fresh water ran out. No person set foot there until 1787.
On Wrangel Island, the last mammoths in the world died about 4,000 years ago. The only dated human camp on the island is about a thousand years younger. Giants can vanish without anyone hunting them: these two islands show that it can happen.
Madagascar
Madagascar, a thousand years ago
People were certainly living on Madagascar 1,350 to 1,100 years ago. Earlier arrivals are argued for: cut marks on elephant-bird bones 10,500 years old, and other evidence from about 4,000 years ago.
Whenever they came, the collapse is dated: elephant birds and pygmy hippos disappeared between about 1,200 and 900 years ago, and some giant lemurs and a hippo lasted a few centuries longer. All seven of the island’s giant mammals are gone, among them a lemur of 160 kg.
New Zealand · about 1280 CE
New Zealand: within two centuries
Polynesians reached New Zealand around 1280. Moa, giant flightless birds up to two metres tall at the back, were gone from the South Island by about 1400–1450, on the dates of 270 bones.
One study estimates there were never more than about 0.01 people per square kilometre. On the globe New Zealand is hatched: its lost giants were birds, and the ochre counts mammals only.
Today
And the giants that survived?
This last picture swaps the possible ranges for the real ones. 154 of the giant species are still alive, but most live in a fraction of the land they could: lions gone from Europe and almost all of India, elephants from most of Africa, bison from most of North America.
The story did not stop. It only slowed down when there were fewer giants left to lose.
Where you live
The giants that would live here
Click the globe or type a town. You get every giant mammal species whose natural range covers that place, drawn to scale beside a person, with when each one was last seen.
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Line the continents up on the day people arrived
Each dot is one extinct giant species, placed at its youngest dated fossil (Andermann et al. 2020). For New Zealand, Mauritius and Madagascar the birds are added from the island studies. The fire band runs from the earliest serious claim of people to the date most researchers accept.
In calendar time the dots are scattered over a hundred thousand years. Slide every row so that zero is the day people arrived, and most of them fall to the right of zero. In the Americas about half fall within 5,000 years of arrival, and on the islands within a few centuries; in Australia and Eurasia they spread over tens of thousands.
Filled dots: last fossil younger than the earliest claim of people. Open dots: older than any claim. Logarithmic scale.
Where
last fossil after the accepted arrival
North America
27 / 31
Australia and New Guinea
15 / 25
Eurasia
28 / 34
South America
25 / 46
Caribbean
1 / 5
Madagascar
4 / 8
The dots that come too early are not proof that those species died before people. A last fossil marks the youngest bone anyone has found and dated, not the last animal alive. In the database the chart uses, Australian species whose last fossil predates the accepted arrival have a median of 5 dated fossil records, against 11 for the rest; in North America, 1 against 14.5. Rare animals vanish from the record early. South America’s chronology is the thinnest: most of its early dots rest on a single dated layer. The chart’s dates come from one database (Andermann et al. 2020) and sometimes differ from the newer studies quoted in the story: its youngest cave bear is 23,900 years old, against 30,500–28,500 in Stuart and Lister (2012).
Was it the end of the Ice Age instead?
The obvious suspect is climate. The extinctions overlap the end of the last Ice Age, when the world warmed by several degrees. So the page asks, for every 96 km square of land: how much did this place warm, from the peak of the Ice Age 21,000 years ago to recent millennia (the reconstruction by Tierney et al. 2020), and what share of its giants is gone?
Take only the places within 30° of the equator that warmed less than 4 °C. The climate changed by about the same amount in all of them. The losses did not:
Median share of giant species lost per 96 km square, squares within 30° of the equator that warmed under 4 °C. Grey: where people, or other humans, lived alongside the animals for hundreds of thousands of years.
Within a landmass, the climate argument does better in some places. Across Eurasia, the more a place warmed, the more giants it lost (rank correlation +0.67); Africa shows the same, more weakly (+0.51). But the most-warmed squares are in the north, where modern humans arrived last and the mammoth steppe disappeared, and this test cannot separate those. In North America and Australia warming tells you almost nothing about which places lost the most (-0.18 and -0.03); South America is weakly positive (+0.28).
Median share lost by how much each square warmed. Bins with fewer than 30 squares are left out.
There is a second argument against climate alone. The same animals had already lived through about ten long ice-age cycles in the last million years, with warmings as fast as this one. Only the last one, the one with people in it, came with a wave of extinctions that took the largest animals first (Svenning et al. 2024; Barnosky et al. 2004).
This repeats, with a finer map and a different climate reconstruction, the test of Sandom et al. (2014), who reached the same conclusion country by country.
The bigger the animal, the more likely it vanished
Climate change has no obvious reason to pick out the largest animals. Hunters do, since a big animal is worth a spear, and slow breeders recover slowly from any loss. Of species between 44 and 100 kg, 41 of 119 are extinct (34%). Above three tonnes, 14 of 16 (88%).
Species extinct, by body mass (PHYLACINE 1.2.1). Africa’s giants mostly survived, at every size up to three tonnes.
In Africa, 1 of 4 species weighing one to three tonnes died out. Everywhere else, 25 of 28.
Where the pattern does not hold
Africa lost giants too, 16 of 82 species, with people present the whole time. Among its best-dated species, the losses fall between 13,000 and 6,000 years ago, which Faith (2014) ties to grasslands changing.
Eurasia lost its giants in a staggered way, over 25,000 years, closely tracking climate swings (Stuart & Lister 2012; Cooper et al. 2015).
St Paul and Wrangel: mammoths died out with no people present; on St Paul, because its fresh water ran out.
Cyprus: the dwarf hippos were “probably already extinct or at least highly endangered” when people arrived (Zazzo et al. 2015).
The Caribbean: on the latest settlement dates, most of the last dated sloth bones come before people.
Arrival dates are argued over: Australia 48,000 or 65,000 years ago; Madagascar 1,350, 4,000 or 10,500. The longer people and giants lived side by side, the weaker “soon after” becomes.
The map rests on an assumption. The ranges of extinct species are drawn where their fossil companions live today, and PHYLACINE’s documentation notes that this treats each extinction as at least partly human-caused. The map shows where these animals would live now; it is not evidence by itself.
Even researchers who blame people warn against the cartoon of hunters sweeping a continent. Koch and Barnosky (2006) call it “an oversimplification”; their own reading is that in many regions people tipped faunas already stressed by climate.
Britain: the mammoths that met people twice
People came and went from Britain with the ice. Human ancestors walked at Happisburgh in Norfolk more than 780,000 years ago. A jaw from Kents Cavern in Devon, 44,200 to 41,500 years old, is among the earliest modern humans in north-west Europe. Then the ice drove everyone out, and people came back to stay only after 14,700 years ago.
The woolly rhinoceros was gone from Britain by about 35,000 years ago. The last British mammoths, the Condover herd in Shropshire, date to 14,500 to 14,000 years ago: within centuries of people returning, and during a sharp warming. In Britain, the people and the climate arrived together.
How this was made
Giants. Land mammals with an average body mass of at least 44 kg in PHYLACINE 1.2.1, which maps the natural range of every mammal alive or extinct in the last 130,000 years on a 96 km grid. Seals, the polar bear and all human species are left out: 329 species, 175 extinct (161 of them before 1500). Status follows the 2016 IUCN list used by PHYLACINE: one of the 175, the scimitar oryx, was then extinct in the wild and has since been returned to Chad, and the wild dromedary is counted although its domestic descendants live on.
Where
species lost
genera lost
Africa
16 / 82 (20%)
12 / 48 (25%)
Eurasia
66 / 147 (45%)
41 / 85 (48%)
North America
50 / 70 (71%)
38 / 54 (70%)
South America
50 / 65 (77%)
41 / 53 (77%)
Australia and New Guinea
25 / 26 (96%)
11 / 12 (92%)
Madagascar
7 / 7 (100%)
4 / 4 (100%)
Caribbean
5 / 5 (100%)
3 / 3 (100%)
When. Each extinct species leaves the globe at its youngest dated fossil, or its last sighting for recent ones, from Andermann et al. (2020); where the young end of a date is unconstrained the old end is used. Four species have no date and are not drawn. Arrival dates per landmass come from the sources below; the globe lights each landmass at once, although people spread across a continent over centuries or millennia.
Climate. Warming is the difference in annual surface air temperature between the Last Glacial Maximum (23,000–19,000 years ago) and the late Holocene (the last 4,000 years) in lgmDA v2.1 (Tierney et al. 2020). Only squares with at least three giant species are used. Squares lie next to each other and are not independent, so no significance tests are given.
Your place. The square under the point you choose. “Gone from here” compares the natural range with today’s range in PHYLACINE. Silhouettes are drawn at each species’ own body mass from a drawing of the same genus or family, so shapes are approximate.
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Natural Earth (1:50m countries and land), public domain; GeoNames towns over 15,000 people (CC BY 4.0); silhouettes from PhyloPic (CC0, CC BY 3.0/4.0; credits below). www.phylopic.org/
Silhouettes: PhyloPic. Mammuthus primigenius: Kamil S. Jaron (CC0); Mammut americanum: artrociraptor (CC0); Palaeoloxodon antiquus: Steven Traver (CC0); Coelodonta antiquitatis: Steven Traver (CC0); Elasmotherium sibiricum: JFstudios (CC0); Megaloceros giganteus: Steven Traver (CC0); Ursus spelaeus: Margot Michaud (CC0); Panthera spelaea: Gabriele Russo (CC0); Bison priscus: Zimices (CC BY 3.0); Bos primigenius: Ivan Iofrida (CC BY 4.0); Equus ferus: Andy Wilson (CC0); Smilodon fatalis: Steven Traver (CC0); Eremotherium laurillardi: Zimices (Julián Bayona) (CC BY 4.0); Doedicurus clavicaudatus: Becky Barnes (CC0); Toxodon platensis: Zimices (CC BY 3.0); Diprotodon optatum: Steven Traver (CC0); Procoptodon goliah: Dan Niel (CC0); Thylacoleo carnifex: Steven Traver (CC0); Genyornis newtoni: Ferran Sayol (CC0); Aepyornis maximus: Robert Cooke (CC0); Dinornis robustus: Ferran Sayol (CC0); Hieraaetus moorei: Andy Wilson (CC0); Raphus cucullatus: Ferran Sayol (CC0); Hydrodamalis gigas: T. Michael Keesey (PDM); Homo neanderthalensis: T. Michael Keesey (CC0); Loxodonta africana: Steven Traver (CC0); Varanus priscus: Rachel T Mason (CC0); Stegodon: Asier Larramendi (CC BY 4.0); Cuvieronius hyodon: Zimices (CC BY 3.0); Notiomastodon platensis: Prehistopia (CC BY 4.0); Homo sapiens: Cagri CEVRIM (CC0).